PARIS — Paris St Germain suffered their first defeat of the season in any competition with Strasbourg ending the runaway leaders' aura of invincibility in Ligue 1 with a 2-1 victory at the Stade de la Meinau on Saturday.

PSG, who had been expected to stretch their 10-point lead with some comfort at a club starting the day in 17th place, were on the end of a major shock as a 65th-minute goal from Stephane Bahoken ended their season-long, 22-match unbeaten sequence.

Later, with the shock of PSG's loss still reverberating, Monaco closed the gap at the top to nine points after Radamel Falcao's early goal earned them a 1-0 victory at home to Angers. Lyon moved second in Ligue 1 after taking advantage of Paris Saint-Germain's slip at Strasbourg with a hard-fought 2-1 win at Caen on Sunday. Maxwel Cornet's tap-in and Mariano Diaz's stunning 12th league goal of the season were enough for Lyon to blast away the blues of their shock home defeat to struggling Lille midweek and move up to 32 points.

Caen pushed Lyon all the way and pulled a goal back through Ivan Santini in stoppage time but stay seventh on 23 points after their defeat. Bruno Genesio's Lyon are nine points behind Neymar's league leaders PSG, who lost their unbeaten record in dramatic >

Marseille, who are fourth on 31 points, will have the chance to overtake Lyon and close the gap between themselves and PSG to seven points when they take on Montpellier later on Sunday.

Lyon came into Sunday's match smarting from their loss to Lille, and took the lead almost immediately, with Cornet lashing home a loose ball from a 10th-minute goalmouth scramble.

Caen kept up the pressure in a scrappy encounter but Diaz made sure of the points nine minutes after the break by crashing a bouncing ball past Caen keeper Remy Vercoutre. The hosts refused to give up and Santini's neat stoppage-time finish gave the home fans hope, but creaking Lyon managed to keep them at bay and just hold on for the win.

Also on Sunday, Saint-Etienne failed to win for a seventh straight French league game as the 10-time champions were held to a 1-1 draw by Nantes, despite a superb goal from Vincent Pajot.

Chasing a first win since October, Saint-Etienne struggled to string passes together in the first half and broke the deadlock against the run of play when Pajot fired a 30-meter (yard) strike into the top left corner. The game was played in a partly closed stadium following incidents that marred Saint-Etienne's defeat to rival Lyon last month. Nantes continued its dominance after the interval and tested Saint-Etienne goalkeeper Stephane Ruffier several times. Emiliano Sala scored a lucky but deserved equalizer in the 61st minute for his fifth goal this season when Leo Lacroix's attempted clearance ricocheted off his leg and head before crossing the line as Ruffier watched powerless.

Nantes pushed hard in the closing stages but substitutes Santy Ngom and Lucas Lima had their efforts saved by Ruffier. Nantes has not won in Saint-Etienne in 17 years and remained fifth in the standings, 14 points behind leader PSG. Saint-Etienne is four points above the relegation zone.

Strasbourg had gone ahead through the inspirational Nuno Da Costa after 13 minutes but normal service seemed to have been resumed when Kylian Mbappe equalized three minutes before halftime.

Despite having to soak up huge pressure after Bahoken again gave Strasbourg the lead, the newly-promoted side survived the onslaught for a famous win, the first over PSG by any team in the league this season after their 13 wins and two draws.

Even having rested their top scorer Edinson Cavani, Marco Verratti and captain Thiago Silva with an eye on Tuesday's Champions League match at Bayern Munich, PSG had the luxury of bringing back the brilliant Mbappe.

Yet they were quickly rocked by the home side's physicality and Dimitri Lienard's free kick saw the excellent Cape Verdean Da Costa take advantage of lax marking to head home his fourth goal of the season.

PSG were let down by poor finishing but Mbappe scored his fifth league goal of the campaign before Javier Pastore hit the post just before halftime.

Yet with Unai Emery's men looking to forge ahead, Strasbourg hit them on the break, goalkeeper Bingourou Kamara's clearance being flicked on by Da Costa into the path of Bahoken, who smashed in a right-foot shot.

From then on, the 25,000-strong crowd must have feared the fairytale would not last against Neymar and company, especially when Cavani was introduced for the last 15 minutes.

When the Uruguayan did get his chance in the eighth of the nine minutes of added time with the Strasbourg goal under siege, the injured Kamara's replacement Alexandre Oukidja blocked Cavani's ricocheted effort after Adrien Rabiot's shot.

Neymar was booked as PSG's frustration grew and he suffered his first defeat since his world-record transfer from Barcelona. Yet it was a wonderful moment for Strasbourg, one of France's iconic clubs now making a stirring comeback after years of struggle when their very future looked in doubt amid a financial crisis.

Falcao, Ligue 1's second most prolific striker this term behind Cavani, headed home his 14th goal of the season for Monaco in just the second minute before the champions made heavy weather of subduing 19th-placed Angers.

Wahbi Khazri and Benjamin Andre struck in the second half for Rennes to earn them a 2-0 win over Amiens, their sixth victory in seven league games, and elevate them to sixth in the table.

Lille moved clear of the relegation zone with Nicolas Pepe's 63rd-minute goal giving them a 1-0 win at home to Toulouse. — Agencies